References - species profiles
click on reference numbers for links to articles and websites
Introduction (return to page)
(1) Rhodin, A., van Dijk, P.P., Iverson, J.B. & Schaffer, H.B. (2009) Turtles of the world, 2010 update: annotated checklist of taxonomy, synonomy, distribution, and conservation status. Pp. 39–84 In: Rhodin, A.G.H., Pritchard, P.C.H., Saumure, R.A., Buhlmann, K.A. & Iverson, J.B. (Eds.), Conservation biology of freshwater turtles and tortoises: a compilation project of the IUCN/SSC Tortoise and Freshwater Turtle Specialist Group.
(Chelonian Research Monographs No. 5).
(2) IUCN red list (accessed 5 Oct 2011)
T. hermanni (return to page)
(1) van Dijk, P.P., Corti, C., Mellado, V.P. & Cheylan, M. (2004) Testudo hermanni. In: IUCN 2011. IUCN Red List of Threatened Species. Version 2011.1. <www.iucnredlist.org>. Downloaded on 28 October 2011
(2) Arnold, E.N., Burton, J.A. & Ovenden, D.W. (1978) Collins field guide: reptiles and amphibians of Britain and Europe. Collins: London
(3) Stubbs, D. (1989) Testudo hermanni, Hermann's tortoise. in I.R.Swingland and M.W.Klemens, (ed.), The conservation biology of tortoises. Occasional Papers of the IUCN Species Survival Commission No. 5.Gland, Switzerland: IUCN. 34-36.
(4) Uwe Fritz, Markus Auer, Albert Bertolero, Marc Cheylan, Tiziano Fattizzo, Anna K. Hundsdörfer, Marcos Martín Sampayo, Joan L. Pretus, Pavel ŠIrokÝ, Michael Wink (2006) A rangewide phylogeography of Hermann's tortoise, Testudo hermanni (Reptilia: Testudines: Testudinidae): implications for taxonomy. Zoologica Scripta,35, 5:531–543
(5) Meek, R. & Innskeep, R. (1980) Aspects of the field biology of a popultion of Hermann's tortoise (Testudo hermanni) in Southern Yugoslavia. British Journal of Herpetology, 6:159-164
(6) Longepierre, S., Hailey, A. & Grenot, C. (2001) Home range area in the tortoise Testudo hermanni in relation to habitat complexity: implicationsfor conservation of biodiversity. Biodiversity and Conservation 10: 1131–1140
(7) Stubbs D and Swingland IR (1985) The ecology of a Mediterranean tortoise (Testudo hermanni): a declining population. Canadian Journal of Zoology 63: 169–180
(8) Rugiero, L. & Luiselli, L. (2006) Ecological modelling of habitat use and the annual activity patterns inan urban population of the tortoise, Testudo hermanni. Italian Journal of Zoology, 73:219-225
(9) Chelazzi, G. & Calzolai, R. (1986) Thermal benefits from familiarity with the environment in a reptile. Oecologica, 68:557-558
(10) Hailey, A. & Willemsen, R.E. (2000) Population density and adult sex ratio of the tortoise Testudo hermanni in Greece: evidence for intrinsic population regulation. Journal of Zoology, 3:325-338
(11) Cheylan, M. (1984) The true status and future if Hermann's tortoise Testudo hermanni robertmertensi WERMUTH 1952 in Western Europe. Amphibia-Reptilia, 5:17-26
(12) Hailey, A (2000) The effects of fire and mechanical habitat destruction on survival of the tortoise Testudo hermanni in northern Greece, Biological Conservation, Volume 92, Issue 3:321–333
(13) Chabanaud, P. (1925) La tortue grecque dans le Midi de la France. Mém. 1er Congrès. Prot. Nat., Paris, 1923, in (11)
(14) Giorgi, F. & Lionello, P. (2008) Climate change projections for the Mediterranean region, Global and Planetary Change, 63:90-104
(15) Fernández-Chacón, A., Bertolero, A., Amengua, A., Tavecchia, G., Homar, V. & Oro, D. (2011) Spatial heterogeneity in the effects of climate change on the population dynamics of a Mediterranean tortoise. Global Change Biology, 17: 3075–3088
T. horsfieldii (return to page)
(1) Fritz, U., Havaš, P. (2007): Checklist of chelonians of the world. Vertebrate Zoology 57: 149-368.
(2) Fritz,U., Auer, M., Chirikova, M.A., Duysebayeva,T.N., Eremchenko, V.K., Kami, H.G., Kashkarov, R.D., Masroor, R., Moodley, Y., Pindrani, A., Široký, P., Hundsdörfer, A.K. (2009) Mitochondrial diversity of the widespread Central Asian steppe tortoise (Testudo horsfieldii Gray, 1844): implications for taxonomy and relocation of confiscated tortoises. Amphibia-Reptilia 30: 245-257
(3) Buhlmann, K. A., Akre, T. S. B., Iverson, J. B., Karapatakis, D., Mittermeier, R. A., Georges, A., Rhodin, A. G. J., van Dijk, P. P., and Gibbons, W. (2009) A global analysis of tortoise and freshwater turtle distributions with identification of priority conservation areas. Chelonian Conservation and Biology, 8: 116-149. In: 18.
(4) Stubbs, D. (1989) Testudo horsfieldii, Horsfield's tortoise. in I.R.Swingland and M.W.Klemens, (ed.), The conservation biology of tortoises. Occasional Papers of the IUCN Species Survival Commission No. 5.Gland, Switzerland: IUCN. 34-36.
(5) Theile, S. (2000) Ranching and breeding Testudo horsfieldii in Uzbekistan,TRAFFIC Europe.
(6) Lagarde F, Bonnet X, Nagy K, Henen B, Corbin J, Naulleau G. (2002) A short spring before a long jump: the ecological challenge to the steppe tortoise (Testudo horsfieldi). Canadian Journal of Zoology. 80:493–502
(7) Highfield, A. C. 1992. The Horsfield's tortoise: Testudo horsfieldi (Gray) 1844 - A brief review of its biology, ecology and captive breeding URL: http://www.tortoisetrust.org/articles/horsfield2.html (accessed 07 Feb 2012).
(8) Lagarde F, Bonnet X, Corbin J, Henen B, Nagy K, Mardonov B, et al. (2003) Foraging behaviour and diet of an ectothermic herbivore: Testudo horsfieldi. Ecography. 26:236–42
(9) Henen, B.T. (2004) Capital and income breeding in two species of desert tortoise. Transactions of the Royal Society of South Africa, 59:65-71
(10) Henen, B.T, Nagy, K.A, Bonnet, X., and Lagarde, F. 2000. Re-productive output of female Central Asian tortoises (Testudo horsfieldi). In Abstracts of the 24th Annual Meeting and Symposium of the Desert Tortoise Council, St. George, Utah, 5-8 March 1999. pp. 44-45. [Abstr.]
(11) Bondarenko, D. A. 1994. Distribution and population density of Horsfield's tortoise in Central Kizil-Kum(Uzbekistan). In Russian. Bulletin MOIP/Moscow Society of Naturalists, 99 (1): 22-27. In: 18.
(12) Grigoryants, A. 2008. Alexandr Grigoryanys. (CITES Management Authority of Uzbekistan) in litt. to CITES Secretariat, 21/07/2008. Information about current status of Horsfield’s tortoise (Agrionemys(Testudo) horsfieldii) in Uzbekistan. In: 18
(13) Devaux, B. 2007. Ouzbekistan – Samarcande tamerlan & les tortues des steppes. La Tortue, 78: 54-75. In 18
(14) Bondarenko, D. A. and Peregontsev, E. A. 2006. Perpectives of study and protection of steppe tortoise inUzbekistan, Second Congrès International Sur La Conservation des Chéloniens, pp. 278-284. In: 18
(15) Mitropolski, O. V. and Kashkarov, R. D. 2000. Information on size of the population of commercial and hunting species of land vertebral animals in the arid region of Uzbekistan. Pp. 21-30 In: Theile, S. 2002. Ranching and breeding Testudo horsfieldii in Uzbekistan - Report to TRAFFIC Europe.
(16) Bozhansky, A. Y. and Polinova, G. V. 2000. Results of quantitative countings of steppe tortoise in the place ofindustrial and potential catching in Uzbekistan. In: Theile, S. 2002. Ranching and breeding Testudohorsfieldii in Uzbekistan - Report to TRAFFIC Europe.
(17) IUCN red list (accessed 7 Feb 2012)
(18) UNEP-WCMC (2010) Review of species selected on the basis of a new or increased export quota in 2010. UNEP-WCMC, Cambridge.
(19) Luxmoore, R., Groombridge, B., and Broad, S. 1988. Significant trade in wildlife: A review of selected species in CITES Appendix II, Vol. 2. Reptiles and invertebrates. IUCN, Gland, Switzerland. 306 pp.
(20) Brusko, Z.K. & Kubykin, R.A. (1982) Horsfield's Tortoise (Agrionemys horsfieldii, Grey 1844) and the way of it's rational utilization in Kazakhstan. Vertebrata Hungarica, 21:55-61. In: Theile, S. (2000) Ranching and breeding Testudo horsfieldii in Uzbekistan,TRAFFIC Europe.
(21) Bondarenko, D. A. and Peregontsev, E. A. 2006. Perpectives of study and protection of steppe tortoise in Uzbekistan, Second Congrès International Sur La Conservation des Chéloniens, pp. 278-284. In:18